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1. Plant survival in the Arctic(after R.M.M. Crawford)
2. Polar areas covering 16.5% of the Earths surface
water availability
extremes and rapid fluctuations in environment (water states)
north and south polar regions differ (geology, energy transport and balance)
3. Arctic and Antarctica Arctic covered by Arctic Ocean (90% multi-year ice), connected to temperate oceans, transport and exchange of heat energy is affected + input of heat from oceanic currents = cyclones, low pressure
Antarctica, a huge continent located at the center of the South polar region, regular movement of air masses, nightless summer + sunless winter, 24 hour radiation in summer brings daily more energy than on the equator
But the sum of energy during the year is low
*Arctic 3 mil.y.a. and Antarctica 25 mil.y.a. => Antarctic sea weeds are better adapted than Arctic macroalgae
4. Arctic tundra
5.5% of the Earths land surface
influenced by man minimally
low temperature
short growing season
limiting factors: thermal input
water
nutrients
5. Phytogeographical division Low Arctic
- shrubs, graminoids
Hight Arctic
- sparse vegetation
- dwarf evergreens, rossete and mat herbs
6. Polarity: wet and dry habitats soil drainage
dry habitats
- dormant earlier
wet habitats
- overwintering buds protected in wet mud
- growth continues until the very last days of summer
7. Temperature problem convenient are sheltered habitats
sun-tracking activity of flowers
flowers parabolic shape (attract pollinators, speed up development and reproduction)
papillose epidermal cells act as lenses
compact rosettes, hairs
8. Photosynthesis high rates of photosynthesis and growth ( temperate plants) but for a shorter time
- exclusively C3 plants
- high activity of enzymes (= metabolic compensation)
- lipids as storage? NO! >cuticle
9. Carefully controlled allocation of carbon
> growth
> reserves
(northern plants more C to underground organs)
northern plants show a high rate of respiration fast mobilization of storage substances
High respiration as the reason for southern limits to the distribution of Arctic plants
10. Plant Nutrients soils 1. skeletal soils of screes, dunes
2. peat (bogs, sedge medows)
limited inputs of nutrients (almost no chemical weathering), only 1 % of nutrients in living biomass
>precipitation
>biological sources (e.g., animal excrements)
the slowest cycling of nutrients
exception: nutrient-rich bird cliffs
11. nitrogen
- wet sites cyanobacteria
- cyanobacteria associated with lichens
- Dryas drummondii N-fixing nodules
phosphorus
- from snowfall and precipitation
(P, Ca content of vegetation may affect population dynamics of lemmings)
12. Reproduction sexual reproduction 10 000x more energy - costly than asexual reproduction
80% of Arctic species perform asexual reproduction (stolons, rhizomes, apomixis)
often sexual and asexual reproduction together
Completion of the sexual reproduction over several seasons (even 7 or 8 years)
13. Altitudinal limits to the survival of trees and shrubs(after R.M.M. Crawford)
14. timberlines: both evergreen conifers and deciduous broad leaved trees
limiting factors: similar to Arctic plants, especially a short growing season (but no permafrost)
15. Tropical timberlines cloud zone: humid climate and stable temperature > little rainfall higher up
timberline not well defined (smaller trees
shrub
mountain desert)\
pachycaul plants woody forms of normally herbaceous plants (Echium, Senecio, Lobelia) protection against fluctuating temperature
Pinus canariensis long needles absorbtion of humidity, dew
16. Temperate timberline 1. abrupt timber-line
- critical stage: seedling establishment
- shade-tolerant trees (Nothofagus) under protection of parental trees
- New Zealand, S America
2. ecotone timber-line
- twisted trees (=Krummholz)
- above snow cover branches exposed to damage
- Europe, N America
- twisted forms genetically fixed,
- special growth regulators (scopulin
)
17. Deciduous X Evergreen Deciduous trees and shrubs:
- investment to new photosynthetic organs every season
- less dangerous during winter
Evergreen trees and shrubs:
- longer time for photosynthesis
- freezing risk during severe winters
18. Altitude of timberline Altitude of timberline decreases by 80m with 1° poleward
S hemisphere: timberline lower then in the N hemisphere (Tyrol 47°N: 2300m X Ands 47°S: 1100m)
tree height is not smaller at the timber-line
19. Temperature effect on tree growth Koppens rule: Tree growth requires a minimal summer warmth represented by the mean daily maximum temperature of 10°C for 1 month
Betula t > 6°C X Pinus July, August t
20. Reproduction Populus tremula vegetative reproduction
Sorbus aucuparia
- lighter seeds but higher growth rates (higher respiration rate)
21. Essential features of mountain plants: plants from lower altitudes - earlier bud break; plants from higher altitudes delayed bud break, but faster life cycle
crucial for survival: to develop sufficient protection and reserves (thick cuticle, carbohydrate storage, etc.)
consequently: long-living trees and shrubs as well as perennial herbs are common in the mountain flora
22. Thank you for your attention! Arctic and mountain plants are admirable!