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Planning. Not all participants can access the field every day as this will impact our study sites and is logistically difficult. In short, participants will spend half-time in the field, and half-time processing and pre-sorting samples in the laboratory.

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Not all participants can access the field every day as this will impact our

study sites and is logistically difficult. In short, participants will spend

half-time in the field, and half-time processing and pre-sorting samples in the


As far as possible the program of each participant as been planned as to

include, out of a sequence of 4 days:

Day 1: Field work: own program

Day 2-3: laboratory

Day 4: field work: own program/individual program/help with other



Tentative planning: scientific activities

Chronologically field work concentrates on crane, bubble, raft and ikos

N.B. Late surveys of FITs = B2: 9 Nov; C3: 10 Nov; R3: 12 Nov, No Ikos


Tentative planning: participants

N.B. The technical staff has been booked as all person being present each day, which is not going to be the case


Field Work

Field work is going to be challenging, no need to hide this. Please be prepared.

Rains will be very frequent: please bring a raincoat or equivalent AND a good

umbrella. Pack all your equipment in plastic bags, use water-resistant paper

for notes, etc. There is no malaria at our field sites or on BCI but chiggers and

ticks are common. Bring insect repellent and medicine against insect bites.

Bring your own climbing gear (harness, jumars, etc.) as to be independent and

efficient. Other useful items: torch and batteries, water bottles (we will use

tap water from BCI for drinking in the field).

In essence: try to be self-sufficient!


Tree-climbing practice

A spot near the crane garita will be designated to test single rope equipment

and to practice tree-climbing, if needed.

Details from Y. Basset, M. Samaniego or B. Corbara.

In doubt, please consult with the professional tree climbers:

T. Aubert, N. Baiben & S. Bechet.


Seasonal replication

Seasonal replication will be discussed between participants during the

first phase of the project. The following is open to discussion:

Vegetation: not needed, but refining measurements at ground plots may be possible

Sticky traps: at crane sites, at least. Needs to leave guide ropes for transects, which could be re-activated also

at other sites.

FITs: same remarks as for sticky traps

Beating: possible at crane sites

Fogging: possible at all sites

Light traps: possible at crane sites

Microarthropods: possible at all sites if guide ropes are left in place

Winkler: not possible at the same sites, but at surrogate sites.

Pitfall traps: possible at all sites

Individual programs: depends on the interest and program


List of focal taxa

Some flexibility may be required as the result of particular sampling programs may be uncertain



Suggested extraction of material










Processing of material

Panamanian students (probably n = 5) will help with the sorting. Priority will be given to programs

generating large amount of specimens, such as fogging, FITs and microarthropods.

Sticky traps: All material identified as far as possible to families and counted.

Focal taxa (n= 22) are extracted and kept in whirl-packs filled with ethanol (2,000 available for

the program). The rest of material is thrown away.

FITs: 33 focal taxa and 3,000 whirl-packs available. Two Panamanian students to help

Beating: 17 focal taxa and 1,500 whirl-packs available

Fogging: 35 focal taxa and 4,000 whirl-packs available. Two Panamanian students to help

Light traps: 18 focal taxa, 2,000 entomological pins and 1,000 whirl-packs available (+ storage boxes)

Microarthropods: 8 focal taxa, 864 vials available (no whirl-packs). One Panamanian student to help

Winkler: 21 focal taxa and 3,000 whirl-packs available

Pitfall traps: 18 focal taxa and 3,000 whirl-packs available

In total: 17,500 whirl-packs and 489 liters ethanol 70 degrees available


Laboratories on BCI

Two main working places have been designated for the IBISCA project on BCI:

1) ‘Wet-dirty’ area: in the Kodak house. Please concentrate all work there which may be

rather wet, dirty or requires handling large samples, etc.

2) ‘Dry-clean’ area: BCI meeting room. For work with stereomicroscopes (n = 11). In addition,

two desktop and one laptop computers will be available for the project in this area

(see databasing,below).

Other working spaces may also be set up in different laboratories. The general BCI computer

room with internet access will be shared with other BCI users.

There will be designated areas for storage of equipment and chemicals.

Databasing: at the end of the first phase, each participant will be required to provide the scientific

coordinators with a simple Excel file detailing the different samples obtained with his/her sampling

program. These files will be needed to coordinates the analyses of the results.

Please think beforehand of the codes you want to use for your samples and how you want to

organize this file.


Collecting, storage and export of specimens

The collecting permit is collective and will be included in your information

envelope (see below). These fees will be paid by the project.

We expect that most of the material will be exported by the participants with

individual export permits, when leaving Panama. However, it may be possible

to store material at the University of Panama. Later exportation of the material

is possible, but more difficult to organize for wet material. Each participant is

required to pay individually for export permits.


Transfer Tocumen airport to BCI

The Hotel Via Espana will host most of the participants when transiting from/to Tocumen airport

and BCI. Booking has been made as indicated in the following page.

Hotel Via Espana: Via Espana (Martin Sosa y Via Espana), Panama City.

Ph. +507 264 0800, Fax +507 264 8802, email

Price single/double rooms: $22/$22-33 (double beds or not).

Transfer airport - Hotel - airport: by taxi/bus organised by Samatour (Ph. 680-8990/646-4660)

The taxi/bus will be dispatched at the airport when you arrive. The driver will wait with a sign

‘IBISCA’. Participants are required to pay the taxi fare ($20, each additional person = $5; i.e.

one person pays $20, 3 people $10 each).

Transfer Hotel - Gamboa - Hotel: Same system organised by Samatour. Cost of fare $20, with additional

person = $5. You need to leave the hotel at least 1 hour 15 before the time departure of the boat.

See next pages for individual bookings and arrival.

Transfer Gamboa - BCI - Gamboa: by boat, free. You need your personal badge to enter the

Dredging Division at Gamboa and access the boat. This badge will be given at your hotel, along with an envelope

with other information.


Arrivals - airport transfers and hotel

As of 11 Sept 2003. If the information is incorrect, please urgently contact Yves!

H. Barrios, Y. Basset, D. Frame, R. Harrison, A. Hernandez, E. Medianero, C. Potvin, D. Roubik, M. Samaniego

& S.J. Wright are based in Panama City and this is irrelevant to them


Barro Colorado Island (BCI)

You are assumed to have signed a form detailing the general conditions of the IBISCA project before

your arrival. This form was sent by email earlier.

Your stay on BCI is also subject to different regulations and recommendations, please see the

documentation sent by e-mail. In particular, washing machines and telephones are available

(for a fee). Meals: 3 meals will be provided per day. However, participants leaving for the field

will be requested to prepare their own breakfast and lunch early in the morning.

Each Thursday, informal seminars are presented on BCI. From our group, F. Hallé, D. Frame

and A. Floren will be presenting such seminars. If you want to present a seminar please contact Yves.

Change: Panama uses US$. If needed, better to change US$ before you leave for Panama, as this

could prove to be time-consuming in the city.

It will be possible to go to Panama City from BCI, by boat and bus (see schedule next page)


Barro Colorado Island:

where to take the boat at Gamboa

In case you need to get there by taxi or bus (in addition, it is possible to get the boat from the City)

Actual loading point:

go here!

Old passenger’s loading point




Rooms - BCI

(as of 4 Sept 03)

Some participants may have been booked longer than indicated on this schedule. When in doubt, please check with Yves


Projected statistical analyses

(data combined from all sampling programs)

This will be discussed by the group, on BCI (see next page). Some suggestions:

Faunal similarity: for each group, between each of the 9 sites (18 sub-sites: upper canopy and

understorey/soil, including vegetation, treated as a taxon or as an explanatory variable.

Correlation between focal taxa, including vegetation, at the 9 (18) sites.

Multivariate analyses, either for particular sampling programs, or as a whole, with vegetation and

canopy openness as explanatory variables for each of the 9 (18) sites.


Meetings and informal discussions

Each day, a short meeting will be held on BCI early in the evening to review

the progress and challenges of the day, and update the daily planning.

It will be possible at that time to call the support crew in Panama City to

review any action needed.

We plan to have a large meeting about mid-way through the project with

all participants, to discuss seasonal replication, processing of material and

statistical analyses.

Other social events and seminars as indicated on next page.


Events and seminars during IBISCA (so far)

  • 22 Sept 2003, 18h00: Inauguration of the ‘Quinzaine scientifique française’ at the University of Panama.
  • 23 Sept 2003, 12h00: Seminar on tree architecture by Francis Hallé, at STRI (Tupper).
  • 25 Sept 2003, 19h15: Seminar on tree architecture by Francis Hallé, at BCI.
  • 1 Oct 2003, 19h00: Public talk (G. Ebersolt, H. Barrios & B. Corbara) about the canopy raft and the project
  • IBISCA (in Spanish) at the University of Panama, for the ‘Quinzaine scientifique française’.
  • 11 Oct 2003: Visit of Smithsonian Institution Secretary Lawrence Small at the IBISCA sites.
  • 12 Oct 2003: Round table with all participants on BCI, to review the progresses, challenges and
  • the future of the IBISCA project.
  • 13 Oct 2003: Visit of Prof. E.O. Wilson at the IBISCA sites.
  • 16 Oct 2003, 19h15: Seminar on comparing arboreal arthropod communities in temperate and tropical forests,
  • by Andreas Floren, at BCI.
  • 20 Oct 2003, 19h15: Seminar on plant-arthropod associations by Dawn Frame, at BCI.
  • 21 Oct 2003, 12h00: Seminar on host specificity, alpha- and beta-diversity of phytophagous beetles in two tropical forests
  • in Panama by Frode Ødegaard

Significance of IBISCA

The major scientific output of the project would represent (i) the first attempt

to compare arthropod species richness in the soil vs. canopy habitats of a tropical

rainforest, including a wide range of taxa and sufficient spatial and temporal

replicates, as well as samples obtained in situ from the canopy; (ii) one of the

first study of beta diversity at a fine scale (a few kilometres) in a lowland

tropical forest, encompassing again a sample size accounting for arthropod

diversity, spatial and seasonal replicates; and (iii) one of the first study of

airborne arthropod fine stratification (a few metres) within the canopy.

The amount of data generated on the vertical stratification and beta-diversity

of 40 focal arthropod taxa phylogenetically and ecologically distant is

likely to have far ranging implications for the conservation of biodiversity

in tropical rainforests.


Participants and duties

Entomologists: Henri-Pierre Aberlenc, Héctor Barrios, Yves Basset, Lukas Cizek, Bruno Corbara,

Gianfranco Curletti, Alain Dejean, Raphael Didham, Laura L. Fagan, Andreas Floren, Rhett Harrison,

Kevin Jordan, Roger Kitching, Maurice Leponce, Enrique Medianero, Frode Ødegaard, Jerome Orivel,

Servio Ribeiro, Yves Roisin, David Roubik, Jürgen Schmidl, Alexey Tishechkin & Neville Winchester

Botanists: Dawn Frame, Francis Hallé, Andres Hernandez, Margaret Lowman, Catherine Potvin,

Mirna Samaniego & Joe Wright

Technical staff: Thierry Aubert, Nouï Baiben, Stephane Bechet, Dany Cleyet-Marrel, Gilles Ebersolt,

Olivier Pascal & Laurent Pyot

Crane operators: José Herrera, Edwin Andrade & Oscar Saldaña

Support crew: Yves Basset, Mirna Samaniego, Patrick Basset, Héctor Barrios & Bruno Corbara

Logistics: Gilles Ebersolt & Mirna Samaniego

Scientific coordination: Yves Basset, Bruno Corbara, Héctor Barrios & Joe Wright



Solvay - Solvin (

Smithsonian Tropical Research Institute (

Global Canopy Programme (

Joe Wright and Yves Basset’s research funds at STRI

Ecoport and University of Florida (

Ambassade de France à Panama (

Note: Ecoport has generously donated $1,000 towards general equipment for the IBISCA project. One requirement is to improve the picture

databank in the Ecoport database (

You can help by donating digital pictures of tropical insect herbivores to Ecoport

(from Panama or elsewhere; pictures of specimens dead or alive). Please contact Yves Basset.


Useful contacts in Panama (+507)

BCI switchboard and fax

Ph.212-8900/8951, Fax 212-8937/8975

IBISCA project, BCI

Ph. 212 8978


Ph. 680-8990 / 646 4660

Yves Basset

Office 603 at STRI (Tupper Building): Ph. 212 8233, Fax 212 8148.

Home 264 9529,


University of Panama (Entomologia), Ph 264 5431,

Mirna Samaniego

Office 523 at STRI, Ph 212 8177,

Orelis Arosemena

Visitor’s office, STRI, Ph 212 8016,

Oris Acevedo

Barro Colorado Island, Ph 212 8901,

Hotel Via Espana

Via Espana (Martin Sosa y Via Espana), Panama City

Ph. 264 0800, Fax 264 8802, email