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第九章 原核生物基因表达的调控 PowerPoint PPT Presentation


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第九章 原核生物基因表达的调控. 基因表达调控就是对基因产物的合成进行控制的机制 . 一个细胞可能仅含有 20 个拷贝乳糖操纵子阻遏蛋白,但是它可能会需要超过 100 000 个拷贝的 EF-Tu 参与蛋白质的合成。 在另外一些情况下,单细胞有机体调控基因表达是为了应答环境条件(例如,温度,渗透压或者是否存在某种营养物质)的改变或者细胞内部的生理状况(例如为细胞分裂做准备)的变化。与细胞适应过程有关的酶或者蛋白质通常是不存在的,只有在需要的时候才被合成。 管家基因与奢侈基因.

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第九章 原核生物基因表达的调控

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  • .

    • 20100 000EF-Tu


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    • mRNA

    • mRNA

    • mRNA


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9.1


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mRNA


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9.1.1

9.1.1.1


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lacA


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lacI


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--D-IPTGIPTG


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lac

catabolite repressioncatabolic activator proteinCAP


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CAPcAMPCAPcAMPcAMP receptor proteinCRPcAMP-CRPRNAPlac35laccAMP-CRPRNARNARNA


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  • araBADAraCaraO2araI1DNAaraBAD

  • AraCaraI1araI2AraCcAMP-CRPRNAPBAD


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  • AraCaraC

  • AraCAraCAraCaraO1araPc


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gal- galEgalTgalKmRNA


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+ ATP 1- + ADP + H+

1- + UDPGlu UDPGal + 1-

UDPGal UDPGlu

+ ATP 1- + ADP + H+


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  • P1P2

  • P2UDPGlu UDPGal

  • OEOIDNADNADNARNA

  • OEP2


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(corepressor)


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regulon


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9.1.1.2

  • 37464630heat shock protein, HSPmolecular chaperones


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  • 32703270


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  • HSP32

    • 3232

    • 32


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E

F


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K

G


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9.1.1.3


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PhoRPhoB


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9.1.2

9.1.2.1

trptrpRtrpRtrptrpR


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trptrptrpEattenuationDNAmRNAattenuatortrp mRNA


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RNA

41234


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1344Utrp2233RNA1234


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9.1.2.2

RNADNA


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9.2

9.2.1 RNA

RNAmRNARNARNAmRNAmRNAmRNA


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RNA

Fur: ferric uptake regulator

FurFurbfrbfrRNA


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9.2.2

E.coli


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autoregulation


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9.2.3 Some mRNA molecules must be cleaved before translation

mRNAadhERNase III


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9.2.3

rRNAtRNAstringent responseppGpppppGppIIIpppGpp


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pppGppRNARNArRNAtRNA

relApppGpprelApppGpprRNAtRNArelArRNAtRNArelArelaxed mutant


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9.3

mRNAmRNAmRNA

7B12TPP()FMN ()SAM (S)


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a); b)


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-6-glmSmRNAGlmS-6--6--6-GlmSGlmS mRNAmRNA


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9.4 DNA

DNASalmonella typhimuriumH1H2


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H1H2H2H1phase variation


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site-specific recombinaseHinHinHinHinHin


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