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Evoluce 7 2013. Glansdorff et al 2008 Biology Direct 2008, 3 :29 . sn -glycerol 3- phosphate fosfolipidy. sn - glycerol 1- phosphate. Archebakterie (Archaea) – doména s unikátním složením membrán. , Eukaryota. and Eukaryota.

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Glansdorffetal2008 Biology Direct 2008, 3:29 


sn-glycerol 3-phosphate

fosfolipidy

sn-glycerol 1-phosphate



Figure 1. membránCommitting steps in the biosynthesis of membrane lipids. The "primary divide" separating Archaea from Bacteria and Eukarya is outlined.

Glansdorffetal 2008 Biology Direct 2008, 3:29 


Luis Villareal: Can viruses make us human? membrán

http://cvr.bio.uci.edu/downloads/03_villa_canvir.pdf

(scifi: Greg Bear: Darwinovo rádio) - zkoušet to nebudu


(7) Archaea arose by reductive evolution under selection for adaptation to high temperatures; this entailed the replacement of sn1,2 ester fatty acid lipid by sn2,3 ether isoprenoid lipids in the cell membrane, without major enzymatic reconversion.

(8) Bacteria arose by reductive evolution; secondary, convergent adaptations to thermophily entailed the formation of a variety of membrane lipids, often with ether bonds, but still with a sn1,2 stereoconfiguration.


(9) LUCA adaptation to high temperatures; this entailed the replacement of wasgeneticallyredundant; thereforethedifferentiallossofparalogous gene copies in different lines ofdescentis a predictedsource ofphylogeneticdiscrepancieswithrespect to the SSU-rRNAtree.


(3) LUCA was a protoeukaryote, with a RNA genome inherited from its progenote ancestor. This RNA LUCA was in a metabolically and morphologically heterogeneous community, constantly shuffling around genetic material. Part of it was phagotrophic. LUCA remained an evolutionary entity, though loosely defined and constantly changing, as long as this promiscuity lasted.

(4) The RNA to DNA transition took place independently in different lineages of this community; the intervention of viruses appears a likely mechanism. This process led to the emergence of the three Domains.


(1) from its progenote ancestor. This RNA LUCA was in a metabolically and morphologically heterogeneous community, constantly shuffling around genetic material. Part of it was phagotrophic. LUCA remained an evolutionary entity, though loosely defined and constantly changing, as long as this promiscuity lasted.Theword "prokaryote" has becomeinadequateandmisleading.

(2) Eukaryoticidiosyncrasiesand a reappraisalofphylogeny do not support thenotionof a prokaryotic LUCA nor ofprokaryoticancestorsoftheeukaryotic cell body. Theorderofbranching in theuniversaltree has no bearing on thecellulararchitectureof LUCA.


Glansdorff from its progenote ancestor. This RNA LUCA was in a metabolically and morphologically heterogeneous community, constantly shuffling around genetic material. Part of it was phagotrophic. LUCA remained an evolutionary entity, though loosely defined and constantly changing, as long as this promiscuity lasted.etal2008 Biology Direct 2008, 3:29 


E.V. Koonin: vývoj buněk z pradávného světa virů from its progenote ancestor. This RNA LUCA was in a metabolically and morphologically heterogeneous community, constantly shuffling around genetic material. Part of it was phagotrophic. LUCA remained an evolutionary entity, though loosely defined and constantly changing, as long as this promiscuity lasted.

Např. Biology Direct 1, 29, 2006

http://www.pubmedcentral.nih.gov/picrender.fcgi?artid=1594570&blobtype=pdf

Biological big bang

Biol. Direct 2,21,2007


Embley & Martin, Nature from its progenote ancestor. This RNA LUCA was in a metabolically and morphologically heterogeneous community, constantly shuffling around genetic material. Part of it was phagotrophic. LUCA remained an evolutionary entity, though loosely defined and constantly changing, as long as this promiscuity lasted.Vol 440|30 March 2006|


  • Bakteriální biosféra from its progenote ancestor. This RNA LUCA was in a metabolically and morphologically heterogeneous community, constantly shuffling around genetic material. Part of it was phagotrophic. LUCA remained an evolutionary entity, though loosely defined and constantly changing, as long as this promiscuity lasted.

  • sdílený genový pool → Bakteriální promiskuita– transdukce, transformace, konjugace

  • sdílené signály → quorumsensing

  • mezibuněčná hmota → konsorcia, povlaky, stromatolity etc.


The greatest surprise resulting from the application of genomics tobacteriology was the extent of genomic variability within many bacterial species.

Bacterial pathogenomics

Mark J. Pallen1 & Brendan W. Wren

NATURE|Vol 449|18 October 2007|


Origins of Highly Mosaic Mycobacteriophage Genomes genomics to

Part of the Che8 genome shown in (A) is

also shared by phage Corndog. The 378 bpshared regions are 100% identical and theirjoint appearance in these phages must bederived from a recent evolutionary event.

Cell, Vol. 113, 171–182, April 18, 2003


Luis Villareal: Can viruses make us human? genomics to

http://cvr.bio.uci.edu/downloads/03_villa_canvir.pdf

(scifi: Greg Bear: Darwinovo rádio) - zkoušet to nebudu

Viry v prostředí: obrovská diverzita (statisíce)

„virioplankton: 106-109 částic/ml

metagenomika

Angly: PLoS Biology 4, e368, 2006


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