[H],
This presentation is the property of its rightful owner.
Sponsored Links
1 / 97

光合作用的过程 PowerPoint PPT Presentation


  • 133 Views
  • Uploaded on
  • Presentation posted in: General

[H],. C 3 化合物还原 : 2 C 3 ( CH 2 O ) 6. 供能. 酶. 光解. 酶. 酶. CO 2 的固定 : CO 2 +C 5 2C 3. 水的光解 : H 2 O 2[H]+1/2 O 2. 光合磷酸化 : ADP+Pi +能量 ATP. ATP. ADP+Pi. 光解. 光合作用的过程. O 2. 2C 3. H 2 O. CO 2. 固 定. 酶. 还 原. [H]. 光能. 吸收. 色素分子. 多种酶.

Download Presentation

光合作用的过程

An Image/Link below is provided (as is) to download presentation

Download Policy: Content on the Website is provided to you AS IS for your information and personal use and may not be sold / licensed / shared on other websites without getting consent from its author.While downloading, if for some reason you are not able to download a presentation, the publisher may have deleted the file from their server.


- - - - - - - - - - - - - - - - - - - - - - - - - - E N D - - - - - - - - - - - - - - - - - - - - - - - - - -

Presentation Transcript


3712721

[H],

C32 C3 CH2O6

CO2: CO2+C5 2C3

H2O 2[H]+1/2 O2

ADP+Pi ATP

ATP

ADP+Pi

O2

2C3

H2O

CO2

[H]

C5

ATP

CH2O

ADP+Pi


3712721

  • :

  • 1.:

  • 2.:;

  • 3.:


3712721

  • 1

  • ATPNADPH2

  • ()10-1510-9 10-1010-4 100101 101102

  • PSPS


3712721


3712721

  • 1012s109s

  • (77K)(2K)

  • 1


3712721

chl a

chla chlb


3712721


3712721

()

  • -----

  • Chl+h 1015s Chl*

8


3712721

  • :

    :

8


3712721

()

1.

2.

3.

4.


3712721

1.

Chl* Chl+ Chl* ChlT + ChlT Chl+


3712721

2.

  • Chl* Chl h

  • ChlT Chl h

  • 1%

10-9s

10-2s


3712721

  • QQ


3712721

3.

  • Chl*1 Chl2 Chl1Chl*2


3712721


3712721

  • ()


3712721


3712721


3712721

()

1.


3712721


3712721


3712721

2.

  • PA h P*A P+A

  • (P+)(A-)


3712721

  • P+ (D)(A1)

  • DP+AA1 D+PAA1


3712721

()PSPS

  • PSPS

  • P700P680


3712721

  • PSPSaP700P680

  • P700680P700nm680nm

9

PS(A)PS(B)700nm(A)682nm(B)


3712721

PSPSPSPSNADP+


3712721

PSPS

  • PS

  • P700A0h P700*0 P700A0

  • PS

  • P680Pheo h P680*Pheo P680+Pheo-


3712721

PSPS


3712721

:

  • :

  • NADP+

  • ATP


3712721

  • (1960)Z

  • Z


3712721

Z

  • (1)PSCyt b/fPS

  • (2)P680P680*P700P700*

10

LHCLHCPSPSMMn


3712721

  • (3)PSNADP+PSPS42H2O1O24H+NADP+

  • (4)PQH+H+H+H+


3712721

()

1.PS

  • PS

    (1)PS

  • PS

PSII


3712721

CP47CP4347 00043 000P680LHCPS


3712721

PS32 00034 000D1D2D1

ZP680PheoQAQBD1D2

D1QBD2QAQ(quinone)(quencher)


3712721

  • QAQBQBQBQA2H+(QH2 )PQPQH2


3712721

(2)

  • (OEC)(M,MSP)PS

  • 1O2H2O 4 e- 4 H

  • CO+2HO* (CHO)+ O2*+ HO


3712721

  • 2060(P. Joliot)

  • (510s300ms)

  • 4OOO41

12

20OO(Joliot1965 )


3712721

  • (B.Kok1970)HO

  • PSHO(S)

  • SPS1e

  • S4e42HO1O(13)

  • SM()MSSSSSSS 1 S44

  • S SS 2 HO 4 eS

13 S

(KokH)


3712721

  • (PQ)PSPSCyt b/f

  • PQ 5%10%

2.


3712721

  • PQ()PSCyt b/f

Cytb6/f


3712721

3.Cytb/f

  • Cytb/f PSPS4CytfCytb Rieske (-)17kD


3712721

  • Cyt b/f PQHPCCyt b/f PQHPC

  • PQH+2PC(Cu) Cyt b/f PQ +2PC(Cu)+ 2H


3712721

Cytb/f MitchellQ

1.PQH22 Cytb6/f FeSRCytf PC

2.PQH2b2H+bbPQ

PQPQ

Q


3712721

4.

  • (PC)Cyt b/fPS


3712721

  • PSPSCyt b/f PC


3712721

5.PS

PSNADP+PCNADP+


3712721

PSLHC


3712721

PSI

A0

A1

FXFAFBFe-S


3712721

14 -

4-4 2-2

FXFAFBPS34-4444-4


3712721

6.-NADP

  • (Fd)-NADP(FNR)

  • Fd2-2

  • FNR 1(FAD)H+FNRFdHNADPNADPH

  • 2Fd+NADP+ H FNR 2Fd+ NADPH


3712721

  • FdPSFd, Fdred

  • FNR

  • Cyt b/f NADPHPQ

  • (Td)


3712721

7.H+

15

4e-2NADPH3ATP8(PSPS4)4e2H2O1 O28H81CO21CO23ATP2NADPH4e-2NADPHATP8H+3ATP


3712721


3712721

()

  • Fd

  • 1.

  • HO PSPQCyt b/fPCPSFdFNR NADP

  • 4e-2H2O1O22NADP+88H+


3712721

2.

  • (1) PS

    PSFdPQCytb/fPCPS

    H2ONADPHH+ATP

  • (2) PS

    P680PheoQAQBCytb559P680

    PS

    P680 Cytb559 Pheo P680

Cytb559


3712721

3.

  • (Mehlers reaction)

  • HOPSPQCytb/fPC PSFd O

  • Fd+ O2 Fd+ O2-

  • (SOD)

  • O2 - + O2 - + 2H2 SOD 2H2O2 + O2


3712721

  • 1954ArnonADPPiNADP+CO2ATPNADPH

ADP PiNADP+

ATPNADPH


3712721

  • ()ADPPiATP


3712721

()

.

  • ATP

  • 2NADP+3ADP3Pi 8h2NADPH3ATPO2+2H+HO

  • ATPNADPH


3712721

.

  • ATP

  • ADP Pi ATP HO

  • ATP


3712721

.

  • ATP

  • HO + ADP Pi ATP + O2-4H+


3712721

()

.--

  • ,

  • (:)


3712721

  • ATP/eP/O

  • ATP/e:ATP

  • P/O1ATP


3712721

2.


Chemiosmotic theory

(chemiosmotic theory)

  • (Mitchell1961)

    • ATP

  • (proton motive force,pmf)


3712721

(1)

  • ATP

  • 1962(Z*)

  • 1963(Jagendorf)Z*H+H+H+ADPPiATP


3712721

-

  • (1963)p44pH8ADPPipHH+

  • HADPPiATPATPpHH+

  • -


3712721

-


3712721

  • pHpHpHH+

  • ATP

  • 1978


3712721

(2)H+

  • eH+,H+(H+) (kJmol-1)

  • H+=H+-H+=2.3RTpH+FE

  • RT(K)FE(V)

  • 25H+ =5.7pH+96.5E

  • F(96.5kJmol1V1)H+/F(V)25pmf=0.059pH+E

pH

E


3712721

ATP


3712721

3.ATPATP

  • ATP(ATPase)ATPCF1-CFoATPATP

  • ATP

  • ATP550 000

  • ATP

  • ATP

H+ transport orientation


3712721

CF1400000,,,5 ::::3:3:1:1:1


3712721

CFo

  • 15

  • CF1CFo

  • CFoCF1ATP

CF1HATP

Paul Boyer


3712721

ATPATP3(tight)(loose)(open)()

ATP-(T)(L)(O)ADPPi ATPATP


3712721

Fo(T)(L)(O)ATP

ADPPi ADPPiATP ATPADPPiATPATPATPATP


3712721

BoyerCF13CF132ATP


3712721

4.

(1)

(2)

(3)ATP

--------


3712721

  • (1)

    (NH2OH)PSDCMUPSQBPQKCNHgPC

    (simazine)(atrazine)(bromacil)(isocil)


3712721

  • (2)

    DNP()CCCP(carbonyl cyanide-3-chlorophenyl hydrazone,-3-)DNH

    H+,


3712721

  • (3)

    ATP(DCCD)(PCMB)CF1CFo(CFo ooligomycin)ATP


3712721

DCMUDBMIB

2,536

DCMUDBMIB


3712721

  • ATPNADPH

  • 1mol ATP50kJ1mol NADPH2mol e0.82V(H2O/O2)0.32V(NADPH) G=nFE=296.5(0.32-0.82)=220kJ

  • , 8mol2molNADPH3molATP680nm8mol(E)

  • E=hNC/8=6.62610s6.023(3.0108msm)8 =1410kJ

  • 8 mol(E) (2molNADPH3molATP)

  • E=220kJ2+50kJ3=590kJ

  • = (/) =E1/E2=590kJ/1410kJ0.42=42%


  • Login