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生物学. 第 15 回 愛は神経とホルモンに乗って. 和田 勝. 「愛」って何?. 定型行動( Fixed action pattern ). たとえば次の例を見てください。. 刺激と反応. 音、光、匂いなど. 外界からの刺激. 個体. 受容器. 中枢神経系. 効果器. 反応. いろいろな行動. ニューロンの数が増える. 脊髄と脳. ニューロンの数が増える. 中枢神経系(脊髄と脳)の中に介在ニューロンによる神経回路が、つくられるようになります。. 特定の神経回路が、定型的行動パターンに対応するようになります(たとえば、歩行運動)。. 定型行動の解発.

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和田 勝

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15


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Fixed action pattern


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Tinbergen


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releaser

key stimuli


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X


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courtship behavior


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oxytocin9


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1909Dale


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1953Vigneaud

SS

Cys-Tyr-Ile-Gln-Asn-Cys-Pro-Leu-GlyNH2


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16CysteinS-S


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G


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10 20 30 40 50 60

1 MEGALAANWS AEAANASAAP PGAEGNRTAG PPRRNEALARVEVAVLCLIL LLALSGNACV

61 LLALRTTRQK HSRLFFFMKHLSIADLVVAV FQVLPQLLWD ITFRFYGPDL LCRLVKYLQV

121 VGMFASTYLLLLMSLDRCLA ICQPLRSLRR RTDRLAVLATWLGCLVASAPQVHIFSLREV

181 ADGVFDCWAV FIQPWGPKAY ITWITLAVYIVPVIVLAACYGLISFKIWQN LRLKTAAAAA

241 AEAPEGAAAG DGGRVALARV SSVKLISKAK IRTVKMTFIIVLAFIVCWTPFFFVQMWSVW

301 DANAPKEASAFIIVMLLASLNSCCNPWIYMLFTGHLFHEL VQRFLCCSAS YLKGRRLGET

361 SASKKSNSSS FVLSHRSSSQ RSCSQPSTA

3p25


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GG

G


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CIP3/DAG

G

C

IP3/DAG

Ca2+


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C

CDAGIP3


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cell membrane

lipid bilayer


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H

H-C-OH

H-C-OH

H-C-OH

H

O

R1-C

O-

O

R2-C

O-

O

R3-C

O-

HO

O H-C-O-C-R1

R2-C-O-C-H

H-C-O-C-R3

H O

+

R


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HO

O H-C-O-C-R1

R2-C-O-C-H O

H-C-O-P-O-X

H O-

phospholipid

phosphatide

X=


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OH

R1-C-O-C-H OCH3

H-C-O-P-O-CH2-CH2-N+-CH3

R2-C-O-C-H O-CH3

OH


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lipid bilayer


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C

IP3Ca2


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OTGq/11IP3/DAG

Ca


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Ca2


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-


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-


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-

neurosecretion


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NHK


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NHK


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Vole

I125

Monogamousprairie volemontane vole

ProNAS 89,5981-5985 (1992)


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prairie vole

montane vole


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amygdala


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NHK


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